A conservation charity has expressed concern over the disappearance of four tagged hen harriers in recent months.
RSPB Scotland said the birds were last located over land managed for grouse shooting when their satellite tags suddenly stopped transmitting.
They were last tracked at locations in Moray, Aberdeenshire and Perthshire.
The Scottish Gamekeepers Association condemned wildlife crime but said there were many other possible explanations for the disappearances.
RSPB Scotland said a chick from a nest in Northumberland whose last known position was on a grouse moor near Grantown-on-Spey in Inverness-shire has not been seen since 16 August.
Two birds tagged on the Mar Lodge Estate in Aberdeenshire this summer were last located near the Aberdeenshire/Moray border on 29 August and near Ballater on 3 September respectively.
One of the birds, Athena, was tagged in Northumberland before disappearing in the Highlands
The fourth missing bird was last recorded on a grouse moor north of Glenalmond in Perthshire on 24 September.
No remains found
Ian Thomson, head of investigations for RSPB Scotland, said: “Each of these missing birds was last known to be on a moor managed for driven grouse shooting before its transmitter suddenly stopped.
“The picture is becoming ever more clear – in almost all cases when a tagged bird dies naturally we are able to recover its remains – if it disappears over a Scottish grouse moor, it’s never seen or heard of again.”
Cathleen Thomas, who manages the charity’s Hen Harrier LIFE project, said: “To have more hen harriers disappear, including three of this year’s youngsters, is devastating for all of us involved in monitoring these hen harrier chicks.”
Hen harriers are among the UK’s rarest birds and are a protected species.
The Scottish Gamekeepers Association has said shooting estates are often unfairly blamed for bird of prey disappearances when there is little supporting evidence.
A spokesman said: “Until the findings of satellite tags are monitored by independent experts or bodies, we will never fully understand what happens when tags lose transmission nor will anyone be any closer to being able to do anything about it.
“There could be many factors at play. Our understanding is that the majority of the seven tagged hen harriers chicks at Mar Lodge this year have died in some circumstance or another, with one tag going off radar for some days before signalling again, so we are not going to speculate on cases.
“Around a quarter of ‘suspicious’ tags studied in SNH’s satellite tagged eagle report lost transmission away from grouse moor areas, including islands, yet the tags themselves were never recovered.”
‘Willing to assist’
Scottish Land and Estates, which represents landowners, said it had only just learned that the hen harriers were missing.
A spokesperson said: “Where a satellite tag stops transmitting, we cannot instantly conclude that the bird has been killed or harmed.
“Land managers across many farms and estates would have been willing to assist the search for these birds had earlier notification been provided.
“If any of these birds have been intentionally killed then that is unlawful and we fully support the full weight of the law being brought to bear on the individuals responsible.”
Anyone with information about the disappearances is asked to contact Police Scotland on 101 or the RSPB’s raptor crime hotline on 0300 999 0101.
Report from BBC Scotland